By Daniel L. Purich

The Advances in Enzymology and similar parts of Molecular Biology Series is without doubt one of the such a lot prestigious within the box, dedicated to the newest developments in molecular biology and enzymology. each one quantity of the sequence includes contributions from prime experts within the box. lower than Dr. Purich's editorship, which all started with quantity seventy two, the sequence has accelerated its insurance to incorporate thematic volumes concentrating on particular study parts, in addition to non-thematic volumes which includes chapters with a extra normal allure.

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By use of pyruvate as a carrier, a fraction waa isolated as the hydrazone. This hydrazone contained but a small per cent of the total fixed radioactive carbon. There was no proof provided, however, that there actually was * However, 4. Proc. NaU. A d . Xci. ,27,476 (1941). 26 WERKMAN AND WOOD any pyruvate formed in the didmilation (none was added apparently). The negative result, therefore, could have been caused either by the absence of an exchange or because there was not enough pyruvate present to give a detectable amount of fixed carbon, even though an exchange did occur.

Krebs and Eggleston (55))reasoning from indirect evidence with pigeon liver and from analogy with bacterial fixation of carbon dioxide (5)) concluded that there is a synthesis of oxalacetic acid from pyruvic acid and carbon dioxide in pigeon liver. The evidence will be considered in greater detail in the section on animal fixation of carbon dioxide, but it is entirely indirect since a net uptake of carbon dioxide was not demonstrated. The proposed function of thiamin in the fixation reaction is baaed on the observation that thiamin, on addition to muscle and liver suspensions from thiamindeficient pigeons, causes an increased dissimilation of pyruvate by liver but not by muscle.

Krebs and Eggleston (44) have shown that the reactions from oxalacetic acid to succinic acid are reversible with the propionic acid bacteria. It has not been shown conclusively, however, that under the conditions of a glycerol fermentation a significant amount of fumarate or succinate is converted to propionate. According to the scheme (Fig. 2) it is eaaential that the reaction proceed as far as a symmetrical molecule (fumarate or succinate) and then reverse itself so that pyruvate may be formed containing fixed carbon.

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